Quarterly report pursuant to Section 13 or 15(d)

Accrued Expenses

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Accrued Expenses
 3 Months Ended
Mar. 31, 2022
Payables and Accruals [Abstract]  
Accrued Expenses Accrued Expenses
Accrued expenses and other current liabilities consisted of the following (in thousands):
  March 31, December 31,
  2022 2021
Accrued accounts payable $ 1,156  $ 2,980 
Collaboration cost-sharing obligation 1,580  1,610 
Research and development fees 5,745  5,682 
Compensation expense 2,454  3,581 
Accrued license obligations 1,500  1,541 
Lease liabilities 1,003  1,049 
Other current liabilities 637  393 
Total $ 14,075  $ 16,836