Exhibit 99.1

 

Joint Filing Agreement

 

The undersigned persons, on February 12, 2018, agree and consent to the joint filing on their behalf of this Schedule 13G in connection with their beneficial ownership of the Common Stock of Pieris Pharmaceuticals, Inc. on December 31, 2017.

 

Tekla Capital Management LLC

 

 

 

 

By

/s/ Laura Woodward

 

Laura Woodward

 

 

 

Daniel R. Omstead

 

 

 

 

By

/s/ Daniel R. Omstead

 

Daniel R. Omstead